A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)

Abstract

Over recent years, intense field work in caves of Portugal has provided new data on the distribution of subterranean Iberian leiodid beetles. Speonemadus algarvensis sp. nov. is described from caves of southern Portugal. The new species is included in the Speonemadus Jeannel, 1922 escaleraigroup (Cholevinae; Anemadini). All species of the S. escalerai-group are revised and S. breuili (Jeannel, 1922) is resurrected as a valid species. A key to identify the species of the S. escalerai-group is provided and the distinctive characters are illustrated. The distribution of all species of the group is mapped with new data together with biogeographic considerations. Key words. Beetles, subterranean, troglobiont, karst, caves, Speonemadus